Field of Study

November 19th, 2025 Associate Professor of Biology Vince Formica investigates beetles to unlock questions of social evolution.

Written by
Tomas Weber

Every summer morning, well before the birds begin to sing, Associate Professor of Biology Vince Formica and a group of students known as the “Beetle Crew” grab their gear from the lab. Climbing onto ATVs, they rumble down an unpaved logging road deep in Appalachia. After arriving at their chosen spot of old-growth forest, they fan out with headlamps, flashlights, and mirrors, peering under fallen logs in search of answers to some of evolutionary biology’s most enduring mysteries.

Whenever they see a forked fungus beetle, the crew gets to work.

“We glue numbers on their backs, and record their codes,” says Formica, who has returned to perform fieldwork at Mountain Lake Biological Station in Southern Virginia every summer for two decades.

Each brown-colored beetle becomes part of a larger puzzle, helping Formica and his team explore how social behaviors — such as cooperation and competition — have shaped the course of evolution.

“The main goal of my research is to understand the evolution of animal societies,” says Formica, for whom the beetles are a potential window into evolutionary processes in general.

“I do that by studying social networks: how individuals are connected, how they interact, and then trying to understand how those individual behaviors lead to big emergent phenomena at the society level.”

Formica wasn’t always the beetle guy. As a doctoral student at Indiana State University, he was interested in sparrows.

Intrigued by how landscapes might influence behavioral differences among birds, he instead noticed that interactions between individuals were shaping their behaviors.

Vince Formica

“One of the ways I do science is, I hand students a data set and give them some papers, and say, ‘What's interesting to you? Go answer that question.’”

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“There’s this thing called the social environment,” he says. “It’s how other individuals are using space. Unlike the physical environment, the social environment is dynamic. It can evolve, too.”

That realization set him on a new course: understanding how the push and pull of social life itself evolves, and how that, in turn, shapes evolution.

“I think the big question is: How does complex social behavior evolve?” he says. “How does aggression, or an individual’s position in its social network, relate to the total structure of society?”

Many researchers tackle these questions theoretically. But Formica wanted empirical answers from the wild. To do that, he needed to find a species with social interactions that could be easily studied. Beetles fit the bill.

“We can go out in the woods, and the populations are all on a single log,” he says. “We can observe a thousand beetles in a single day. They can live for six years — so I’ve known some of these beetles for longer than some of my friends.”

He was also looking for a species that he could easily bring undergraduates to study.

“I knew I wanted to be at a liberal arts school,” says Formica, who joined Swarthmore in 2012.

His own experience of field research as an undergraduate at St. Mary’s College of Maryland was seminal.

“It was literally life changing for me,” says Formica. “Giving that opportunity to Swarthmore students drives what I do.”

Field biology still suffers from a lack of diversity, and Formica hopes to give his students, who hail from a wide range of backgrounds, an appreciation of the beauty and complexity of life in the wild — as well as their first introduction to life without creature comforts.

“Some students don’t even own a pair of hiking boots,” he says. “But they move to this remote field study on top of a mountain at 3,000 feet in Southern Virginia without cell signal. And then they’re out in the dark, braving the rattlesnakes and bears.”

Years of this grueling, but invariably entertaining, fieldwork have yielded valuable insights. To name but one: Years of close observation and data collection revealed a male beetle’s chances of reproductive success hinges not only on its own horn size, but also on the horn dimensions of its chosen male companions. Large-horned males frequently associate with smaller-horned individuals, boosting their chances of success.

The takeaway? Individual success is not just about individual traits. It’s also about how those traits interact with the surrounding community.

“We’ve discovered that the social environment is really important to the success of individuals in a population,” he says.

Formica also encourages members of the Beetle Crew to go off and chase answers to the questions that fascinate them.

“One of the ways I do science is, I hand students a data set and give them some papers, and say, ‘What's interesting to you? Go answer that question.’”

Thanks to these independent student projects, we now know that male beetles consistently prefer larger and more fecund females, and that beetles tend to have more offspring the older they get.

But despite decades in the field, Formica is still struck by the weight of what he doesn’t know.

“I’ve been doing this for 20 years,” he says. “I’d have imagined I knew everything I needed to know about this system. But there’s just more, there’s always more.”

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